Col Dmytro Kozyura agreed to share state secrets and systematically disclosed classified information, prosecutors say.
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Hundreds are still feared trapped under the rubble, as families desperate for news.
Bolton faces a prison sentence of up to five years and has agreed to pay $2.25m in fine, prosecutors say.
US Central Command says it has struck missile and drone storage facilities and coastal radar positions.
Critics say the new reading requirements infringe on religious freedoms and blur the separation of church and state.
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